[A]nalysis of the chemical composition of their bones reveals that they were obligate carnivores.

In addition to being the obligate food source for monarch caterpillars, milkweeds also provide abundant nectar for the adult butterflies.

In some languages such signaling is optional, whereas in others it is obligate.

Aquatic sites constitute obligate habitat for some species, and are critical breeding habitat for species with complex life cycles involving aquatic egg or larval development.

Unlike for phagotrophic flagellates, bacteria serve as a facultative rather than an obligate food source for crustacean zooplankton.

Light is the obligate energy source for the phototrophic microbes constructing these benthic mats

Obligate

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